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Génétique Quantitative et Évolution - Le Moulon



Directeur de Recherche, INRAE

Diversity management for agroecology


01 69 15 68 55



  • Génétique Quantitative et Évolution - Le Moulon
  • Université Paris-Saclay, INRAE, CNRS, AgroParisTech
  • Ferme du Moulon
  • F-91190 Gif-sur-Yvette

Education and positions

  • Senior Researcher (“Directeur de Recherche”), Institut National de la Recherche Agronomique (INRA)
  • Head of DEAP team (Population Diversity, Evolution and Adaptation) at GQE-Le Moulon
  • 1998: PhD thesis INA-PG
  • 1993: ingénieur Agronome and MS in Plant Breeding INA-PG (present AgroParisTech)/Univ. Paris-Sud

Research interests

My current research aims at:

  • Understanding how the management of intra-specific diversity in crops can contribute to a better performance of crops, and more specifically study the effect of plant to plant interactions in wheat cultivar mixtures, through experimental approaches as well as development of models coupling ecophysiology and genetics (Wheatamix - WALTer);
  • Developing appropriate models in quantitative genetics to study the mixing ability of genotypes and contribute to the development of original breeding schemes for plant teams, both within and between different crop species;
  • Mobilizing participatory approaches to co-design mixtures delivering appropriate services from farmers to end-users;
  • Studying the impact of the managementof disease resistances on the evolution of virulence of pathogen populations, focusing on wheat cultivars deployment in Europe and the selective pressure produced on rust fungal populations (RustWatch).

Main ongoing / recent external collaborators

  • France Sébastien Barot (IeeS, Paris) ; Emmanuelle Porcher (MNHN, Paris) ; Arnaud Gauffreteau et Marie-Hélène Jeuffroy (Agronomie Grignon) ; Vincent Allard (GDEC, Clermont) ; Rémi Perronne (INRA, IGEPP, Rennes) ; C. Pope et M. Leconte (BIOGER, Thiverval-Grignon) ; Pierre Barbillon et Christophe Ambroise (MIA- Paris) ; Christian Fournier et Christophe Pradal (LEPSE, AGAP, Montpellier) ; Nathalie Moutiers et Alain Baranger (IGEPP, Rennes) ; Isabelle Litrico et Cyril Firmat (URP3F, Lusignan) ; Sophie Bouchet (GDEC), Jean-Noël Aubertot (AGIR) ; Valérie Cadot (GEVES, Angers) ; Matthieu Bogard et Philippe du Cheyron (ARVALIS, Paris).

  • International Benedikt Haugh, Pierre Hohmann, Monika Mesmer (Fibl, SW) ; Maria Finkh (U. Kassel, DE) ; Mogens Hovmoller (Aarhus U., DK) ; Ian Mackay (NIAB, UK) ; Sajid Ali (U. Peshawar, PK).

Students training

(Co-)supervision of 7 PhD Student:

Mamadou Mboup (2005-2008), Bahri Bochra (2006-2009), Sajid Ali (2010-2013), Abdul Reyman Khan (2010-2013), Stéphanie Thépot (2010-2014), Christophe Lecarpentier (2012-2015), Emma Forst (2014-2017), Adrien Speck (2017-2020), Emmanuelle Blanc (2017-2020), Benedikt Haug (2018-2021).

Research management

  • Elected member of CCSU 64-69 (Université Paris-Sud) from 2010.
  • Member of CSS EGBIP (INRA), 2011-2014.
  • Member of the section 29 of the National Comitee for Scientific Research (CoNRS) from 2012.
  • 2010-2013: Director of Federal Institute for Research “the Plant and its environment” (IFR87, closed since December 2013)


  • Haug B., Messmer MM., Enjalbert J. , Goldringer I., Forst E., Flutre T., Mary-Huard T., Hohmann P.. (2021) Advances in Breeding for Mixed Cropping – Incomplete Factorials and the Producer/Associate Concept. Frontiers in Plant Science,
  • Forst E., Enjalbert J. , Allard V., Ambroise C., Krissaane I., Mary-Huard T., Robin S., Goldringer I.. (2019) A generalized statistical framework to assess mixing ability from incomplete mixing designs using binary or higher order variety mixtures and application to wheat. Field Crops Research, (242) 107571
  • Lecarpentier C., Barillot R., Blanc E., Abichou M., Goldringer I., Barbillon P., Enjalbert J. , Andrieu B.. (2019) WALTer: a three-dimensional wheat model to study competition for light through the prediction of tillering dynamics. Ann Bot, 6 (123) 961-975
  • Borg J., Kiær LP., Lecarpentier C., Goldringer I., Gauffreteau A., Saint-Jean S., Barot S., Enjalbert J. . (2018) Unfolding the potential of wheat cultivar mixtures: A meta-analysis perspective and identification of knowledge gaps. Field Crops Res, (221) 298-313
  • Barot S., Allard V., Cantarel A., Enjalbert J. , Gauffreteau A., Goldringer I., Lata JC., Le Roux X., Niboyet A., Porcher E.. (2017) Designing mixtures of varieties for multifunctional agriculture with the help of ecology. A review. Agron. Sustain. Dev., 2 (37) 13
  • Perronne R., Diguet S., Vallavieille-Pope C., Leconte M., Enjalbert J. . (2017) A framework to characterize the commercial life cycle of crop varieties: Application to the case study of the influence of yellow rust epidemics on French bread wheat varieties. Field Crops Research, (209) 159-167
  • Enjalbert J. , Allard V., Andrieu B., Barot S., Borg J., Cantarel A., Feret M., Cervek C., Coleno FC., Pope De Vallavieille C., Descoureaux D., Dubs F., Galic N., Gauffreteau A., Gilet JD., Goldringer I., Hannachi M., Houivet G., Pin S., Jeuffroy MH., Kerbiriou C., Labarthe P., LATA JC., LeCarpentier C., Lejars L., Lemain B., Lemarié S., Leny F., Le Roux X., Le Viol I., Montagnier C., NIBOYET A., Omon B., Piaud S., Poly F., Pommier T., Porcher E., Saint-Jean S., Salmon S., Tropée D., Vidal T.. (2016) Increasing species richness and genetic diversity in agriculture: results of the Wheatamix project. , 40 p.
  • Thépot S., Restoux G., Goldringer I., Hospital F., Gouache D., Mackay I., Enjalbert J. . (2015) Efficiently Tracking Selection in a Multiparental Population: The Case of Earliness in Wheat. Genetics, 2 (199) 609-623
  • Thépot S., Restoux G., Hospital F., Gouache D., Mackay I., Goldringer I., Enjalbert J. , Ogihara Y., Takumi S., Handa H.. (2015) Interest of a Multiparental and Outcrossing Wheat Population for Fine Mapping. , 331-8
  • Mboup M., Bahri B., Leconte M., De Vallavieille-Pope C., Kaltz O., Enjalbert J. . (2012) Genetic structure and local adaptation of European wheat yellow rust populations: the role of temperature-specific adaptation. Evol. Appl., 4 (5) 341-352